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Showing The Origin of Species Illustrated (The Illustrated Origin of Species) order by subject, title, ordinal limit 116, 4 (4 of 475).
# subject title description
117
04 - Natural Selection
04-04 - Its Power at All Ages and on Both Sexes
As we see that those variations which, under domestication, appear at any particular period of life, tend to reappear in the offspring at the same period;- for instance, in the shape, size, and flavour of the seeds of the many varieties of our culinary and agricultural plants; in the caterpillar and cocoon stages of the varieties of the silk-worm; in the eggs of poultry, and in the colour of the down of their chickens; in the horns of our sheep and cattle when nearly adult;- so in a state of nature natural selection will be enabled to act on and modify organic beings at any age, by the accumulation of variations profitable at that age, and by their inheritance at a corresponding age.
Full Size

silkworm
silkworm

egg
egg

poultry
poultry

chicken
chicken

sheep
sheep

cattle
cattle


If it profit a plant to have its seeds more and more widely disseminated by the wind, I can see no greater difficulty in this being effected through natural selection, than in the cotton-planter increasing and improving by selection the down in the pods on his cotton-trees.

cotton
cotton


Natural selection may modify and adapt the larva of an insect to a score of contingencies, wholly different from those which concern the mature insect; and these modifications may affect, through correlation, the structure of the adult.

larva
larva

insect
insect


So, conversely, modifications in the adult may affect the structure of the larva; but in all cases natural selection will ensure that they shall not be injurious: for if they were so, the species would become extinct.
118
04 - Natural Selection
04-04 - Its Power at All Ages and on Both Sexes
Natural selection will modify the structure of the young in relation to the parent, and of the parent in relation to the young.

In social animals it will adapt the structure of each individual for the benefit of the whole community, if the community profits by the selected change.

What natural selection cannot do, is to modify the structure of one species, without giving it any advantage, for the good of another species; and though statements to this effect may be found in works of natural history, I cannot find one case which will bear investigation.

A structure used only once in an animal's life, if of high importance to it, might be modified to any extent by natural selection; for instance, the great jaws possessed by certain insects, used exclusively for opening the cocoon- or the hard tip to the beak of unhatched birds, used for breaking the egg.

insect
insect

bird
bird

beak
beak

egg
egg


It has been asserted, that of the best short-beaked tumbler-pigeons a greater number perish in the egg than are able to get out of it; so that fanciers assist in the act of hatching.

Tumbler Pigeon
Tumbler Pigeon


Now if nature had to make the beak of a full-grown pigeon very short for the bird's own advantage, the process of modification would be very slow, and there would be simultaneously the most rigorous selection of all the young birds within the egg, which had the most powerful and hardest beaks, for all with weak beaks would inevitably perish; or, more delicate and more easily broken shells might be selected, the thickness of the shell being known to vary like every other structure.

It may be well here to remark that with all beings there must be much fortuitous destruction, which can have little or no influence on the course of natural selection.

For instance a vast number of eggs or seeds are annually devoured, and these could be modified through natural selection only if they varied in some manner which protected them from their enemies.

egg
egg

seeds
seeds


Yet many of these eggs or seeds would perhaps, if not destroyed, have yielded individuals better adapted to their conditions of life than any of these which happened to survive.

So again a vast number of mature animals and plants, whether or not they be the best adapted to their conditions, must be annually destroyed by accidental causes, which would not be in the least degree mitigated by certain changes of structure or constitution which would in other ways be beneficial to the species.

But let the destruction of the adults be ever so heavy, if the number which can exist in any district be not wholly kept down by such causes,- or again let the destruction of eggs or seeds be so great that only a hundredth or a thousandth part are developed,- yet of those which do survive, the best adapted individuals, supposing that there is any variability in favourable direction, will tend to propagate their kind in larger numbers than the less well adapted.

If the numbers be wholly kept down by the causes just indicated, as will often have been the case, natural selection will be powerless in certain beneficial directions; but this is no valid objection to its efficiency at other times and in other ways; for we are far from having any reason to suppose that many species ever undergo modification and improvement at the same time in the same area.
119
04 - Natural Selection
04-05 - Sexual Selection
Inasmuch as peculiarities often appear under domestication in one sex and become hereditarily attached to that sex, so no doubt it will be under nature.

Thus it is rendered possible for the two sexes to be modified through natural selection in relation to different habits of life, as is sometimes the case; or for one sex to be modified in relation to the other sex, as commonly occurs.

This leads me to say a few words on what I have called Sexual Selection. This form of selection depends, not on a struggle for existence in relation to other organic beings or to external conditions, but on a struggle between the individuals of one sex, generally the males, for the possession of the other sex.

The result is not death to the unsuccessful competitor, but few or no offspring. Sexual selection is, therefore, less rigorous than natural selection.

Generally, the most vigorous males, those which are best fitted for their places in nature, will leave most progeny.

But in many cases, victory depends not so much on general vigor, as on having special weapons, confined to the male sex.

A hornless stag or spurless cock would have a poor chance of leaving numerous offspring.

stag
stag

cock
cock

spurs
spurs


Sexual selection, by always allowing the victor to breed, might surely give indomitable courage, length to the spur, and strength to the wing to strike in the spurred leg, in nearly the same manner as does the brutal cockfighter by the careful selection of his best cocks.

cockfight
cockfight


How low in the scale of nature the law of battle descends, I know not; male alligators have been described as fighting, bellowing, and whirling round, like Indians in a war-dance, for the possession of the females; male salmons have been observed fighting all day long; male stagbeetles sometimes bear wounds from the huge mandibles of other males; the males of certain hymenopterous insects have been frequently seen by that inimitable observer M. Fabre, fighting for a particular female who sits by, an apparently unconcerned beholder of the struggle, and then retires with the conqueror.

alligator
alligator

crocodile
crocodile

Stag Beetle
Stag Beetle

Indian
Indian


The war is, perhaps, severest between the males of polygamous animals, and these seem oftenest provided with special weapons.

The males of carnivorous animals are already well armed; though to them and to others, special means of defence may be given through means of sexual selection, as the mane of the lion, and the hooked jaw to the male salmon; for the shield may be as important for victory, as the sword or spear.

mane
mane

salmon
salmon

sword
sword

spear
spear
120
04 - Natural Selection
04-05 - Sexual Selection
Amongst birds, the contest is often of a more peaceful character. All those who have attended to the subject, believe that there is the severest rivalry between the males of many species to attract, by singing, the females.

bird
bird


The rock-thrush of Guiana, birds of paradise, and some others, congregate; and successive males display with the most elaborate care, and show off in the best manner, their gorgeous plumage; they likewise perform strange antics before the females, which, standing by as spectators, at last choose the most attractive partner.

Rock Thrush
Rock Thrush

Rock Thrush (female)
Rock Thrush (female)

Bird of Paradise
Bird of Paradise

Bird of Paradise (flower)
Bird of Paradise (flower)


Those who have closely attended to birds in confinement well know that they often take individual preferences and dislikes: thus Sir R. Heron has described how a pied peacock was eminently attractive to all his hen birds.

peacock
peacock


I cannot here enter on the necessary details; but if man can in a short time give beauty and an elegant carriage to his bantams, according to his standard of beauty, I can see no good reason to doubt that female birds, by selecting, during thousands of generations, the most melodious or beautiful males, according to their standard of beauty, might produce a marked effect.

Some well-known laws, with respect to the plumage of male and female birds, in comparison with the plumage of the young, can partly be explained through the action of sexual selection on variations occurring at different ages, and transmitted to the males alone or to both sexes at corresponding ages; but I have not space here to enter on this subject.

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Statistics and Drill Down Data Mining
subject #
01 - Variations Under Domestication 54 54
02 - Variations Under Nature 23 77
03 - Struggle for Existence 30 107
04 - Natural Selection 105 212
05 - Laws of Variation 48 260
06 - Difficutiles in Theory 74 334
07 - Instinct 13 347
08 - Hybridism 9 356
09 - On the Imperfection of the Geological Record 7 363
10 - On The Geological Succession of Organic Beings 10 373
11 - Geographical Distribution 7 380
12 - Geographical Distribution -- continued 23 403
13 - Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Or 34 437
14 - Recapitulation and Conclusion 38 475
title #
01-01 - Causes of Variability 5 5
01-02 - Effects of Habit 2 7
01-03 - correlation of Growth 2 9
01-04 - Inheritance 4 13
01-05 -Character of Domestic Varieties 2 15
01-06 - Difficulty of distinguishing between Varieties and Species 2 17
01-07 - Origin of Domestic Varieties from one or more Species 7 24
01-08 - Breeds of the Domestic Pigeons, their Differences and Origin 9 33
01-09 - Principles of Selection anciently followed, and their Effects 6 39
01-10 - Methodical and Unconscious Selection 5 44
01-11 - Unknown Origin of our Domestic Productions 5 49
01-12 - Circumstances favourable to Man's Power of Selection 3 52
01-13 - Summary 2 54
02-01 - Variability 2 56
02-02 - Individual Differences 2 58
02-03 - Doubtful Species 10 68
02-04 - Wide-ranging, much diffused, and common Species vary most 3 71
02-05 - Species of the Larger Genera in each Country vary more frequently than the Species of the Smaller Genera 2 73
02-06 - Many of the Species included within the Larger Genera resemble Varieties in being very closely, but unequally, related to each other, and in having restricted ranges 2 75
02-07 - Summary 2 77
03-01 - Bears on Natural Selection 2 79
03-02 - The Term, Struggle for Existence, used in a large sense 2 81
03-03 - Geometrical Ratio of Increase 2 83
03-04 - Rapid Increase of naturalised Animals and Plants 3 86
03-05 - Nature of the Checks to Increase 2 88
03-06 - Competition Universal 2 90
03-07 - Effects of Climate 2 92
03-08 - Protection from the Number of Individuals 2 94
03-09 - Complex Relations of all Animals and Plants Throughout Nature 5 99
03-10 - Struggle for Life most severe between Individuals and Varieties of the same Species 2 101
03-11 - The Relation of Organism to Organism the Most Important of All Relations 4 105
03-12 - Summary 2 107
04-01 - Natural Selection 5 112
04-02 - Its Power Compared with Man's Selection 2 114
04-03 - Its Power on Characters of Trifling Importance 2 116
04-04 - Its Power at All Ages and on Both Sexes 2 118
04-05 - Sexual Selection 3 121
04-06 - On the generality of Intercross Between Individuals of the Same Species 9 130
04-07 - Illustrations of the Action of Natural Selection: 10 140
04-08 - On the Intercrossing of Individuals 8 148
04-09 - Circumstances favourable for the production of new forms through Natural Selection 12 160
04-10 - Extinction caused by Natural Selection 3 163
04-11 - Divergence of Character 26 189
04-12 - On the Degree to which Organisation tends to advance 11 200
04-13 - Convergence of Character 8 208
04-14 - Summary of Chapter 4 212
05-01 - Effects of External Conditions 2 214
05-02 - Use and Disuse of Parts, combined with Natural Selection, Organs of Flight and Vision 7 221
05-03 - Acclimatisation 4 225
05-04 - Correlation of Growth 5 230
05-05 - Compensation and Economy of Growth 2 232
05-06 - False Correlation 2 234
05-07 - Multiple, Rudimentary, and Lowly-organised Structures are Variable 2 236
05-08 - Parts Developed in an Unusual Manner are Highly Variable 5 241
05-09 - Specific Characters more Variable than Generic Characters 2 243
05-10 - Secondary Sexual Characters Variable 3 246
05-11 - Species of the Same Genus Vary in an Analogous Manner 2 248
05-12 - Reversion to Long Lost Characters 10 258
05-13 - Summary 2 260
06-01 - Difficulties on the Theory of Descent with Modification 5 265
06-02 - Transitions 2 267
06-03 - Absence or Rarity of Transitional Varieties 10 277
06-04 - Transitions in Habits of Life 7 284
06-05 - Diversified Habits in the Same Species 2 286
06-06 - Species with Habits Widely Diffferent from those of their Allies 3 289
06-07 - Organs of extreme Perfection 5 294
06-08 - Means of Transition 6 300
06-09 - Cases of Difficulty 5 305
06-10 - Natura Non Facit Saltum 2 307
06-11 - Organs of Small Importance 6 313
06-12 - Organs not in all Cases Absolutely Perfect 13 326
06-13 - Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by the Theory of Natural Selection 8 334
07-01 - Instincts comparable with habits, but different in their origin 2 336
07-02 - Instincts Graduated 2 338
07-03 - Aphides and ants 1 339
07-04 - Instincts variable 1 340
07-05 - Domestic instincts, their origin 1 341
07-06 - Natural instincts of the cuckoo, ostrich, and parasitic bees 1 342
07-07 - Slave-making ants 1 343
07-08 - Hive-bee, its cell-making instinct 1 344
07-09 - Difficulties on the theory of the Natural Selection of instincts 1 345
07-10 - Neuter or sterile insects 1 346
07-11 - Summary 1 347
08-01 - Distinction between the sterility of first crosses and of hybrids 1 348
08-02 - Sterility various in degree, not universal, affected by close interbreeding, removed by domestication 1 349
08-03 - Laws governing the sterility of hybrids 1 350
08-04 - Sterility not a special endowment, but incidental on other differences 1 351
08-05 - Causes of the sterility of first crosses and of hybrids 1 352
08-06 - Parallelism between the effects of changed conditions of life and crossing 1 353
08-07 - Fertility of varieties when crossed and of their mongrel offspring not universal 1 354
08-08 - Hybrids and mongrels compared independently of their fertility 1 355
08-09 - Summary 1 356
09-01 -On the absence of intermediate varieties at the present day 1 357
09-02 - On the nature of extinct intermediate varieties; on their number 1 358
09-03 - On the vast lapse of time, as inferred from the rate of deposition and of denudation 1 359
09-04 - On the poorness of our palaeontological collections 1 360
09-05 - On the intermittence of geological formations 1 361
09-06 - On the absence of intermediate varieties in any one formation 1 362
09-07 - On their sudden appearance in the lowest known fossiliferous strata 1 363
10-01 - On the slow and successive appearance of new species 1 364
10-02 - On their different rates of change 1 365
10-03 - Species once lost do not reappear 1 366
10-04 - Groups of species follow the same general rules in their appearance and disappearance as do single species 1 367
10-05 - On Extinction 1 368
10-06 - On simultaneous changes in the forms of life throughout the world 1 369
10-07 - On the affinities of extinct species to each other and to living species 1 370
10-08 - On the state of development of ancient forms 1 371
10-09 - On the succession of the same types within the same areas 1 372
10-10 - Summary of preceding and present chapters 1 373
11-01 - Present distribution cannot be accounted for by differences in physical conditions 1 374
11-02 - Importance of barriers 1 375
11-03 - Affinity of the productions of the same continent 1 376
11-04 - Centres of creation 1 377
11-05 - Means of dispersal, by changes of climate and of the level of the land, and by occasional means 2 379
11-06 - Dispersal during the Glacial period co-extensive with the world 1 380
12-10 - Distribution of fresh-water productions 1 381
12-20 - On the inhabitants of oceanic islands 1 382
12-30 - Absence of Batrachians and of terrestrial Mammals 4 386
12-40 - On the relations of the inhabitants of islands to those of the nearest mainland 3 389
12-50 - On colonisation from the nearest source with subsequent modification 10 399
12-60 - Summary of the last and present chapters 4 403
13-01 - CLASSIFICATION, groups subordinate to groups 1 404
13-02 - Natural system 1 405
13-03 - Rules and difficulties in classification, explained on the theory of descent with modification 13 418
13-04 - Classification of varieties 1 419
13-05 - Descent always used in classification 6 425
13-06 - Analogical or adaptive characters 2 427
13-07 - Affinities, general, complex and radiating 2 429
13-08 - Extinction separates and defines groups 4 433
13-09 - MORPHOLOGY, between members of the same class, between parts of the same individual 1 434
13-10 - EMBRYOLOGY, laws of, explained by variations not supervening at an early age, and being inherited at a corresponding age 1 435
13-11 - RUDIMENTARY ORGANS; their origin explained 1 436
13-12 - Summary 1 437
14-01 - Recapitulation of the difficulties on the theory of Natural Selection 7 444
14-02 - Recapitulation of the general and special circumstances in its favour 10 454
14-03 - Causes of the general belief in the immutability of species 13 467
14-04 - How far the theory of natural selection may be extended 1 468
14-05 - Effects of its adoption on the study of Natural history 5 473
14-06 - Concluding remarks 2 475
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